Integrating epigenomic features reveals principles of chromatin-state organization Joseph Martini1*, Ryan A Williams2346*, Rebecca G Smith2345, Yan Zhou2346# and Yu Liu2346# 1Lewis Katz School of Medicine, Temple University, Philadelphia, PA 19122 2Cancer Epigenetics Institute, Fox Chase Cancer Center, Philadelphia, PA 19111, USA 3Nuclear Dynamics and Cancer Program, Fox Chase Cancer Center, Philadelphia, PA 19111, USA 4Institute for Cancer Research, Fox Chase Cancer Center,